The four chicks which hatched as part of a rare osprey love triangle in the Borders have died.
Forestry and Land Scotland (FLS) had captured the unusual arrangement on cameras set up as part of the Tweed Valley Osprey Project (TVOP) at Glentress near Peebles.
Two female birds and one male had been breeding in what initially appeared to be a "tolerant" relationship.
However, after the male bird left the nest, the two females struggled to provide enough food for the chicks which ultimately failed to survive.
The unusual breeding situation was revealed last month when the relationship between the birds was "looking good".
They worked together to incubate the four eggs in the nest and the chicks began to hatch on 28 May.
But by that stage the male bird – named Newboy – had abandoned the nest, leaving the two females – F2 and Mrs O – to provide for the chicks.
Project co-ordinator Diane Bennett said that process had started out quite well.